Reflects improved gnomAD gene_symbol fallback (+3,471 genes). Adds dual ranking (≥3 vs ≥4 layers) to address normalization inflation. New high-priority candidates: ARL3 (52% cilia literature), HDAC6 (known ciliogenesis regulator). HIGH tier: 82 genes. Co-Authored-By: Claude Opus 4.6 <noreply@anthropic.com>